The best thing about Auto Orienting 3D layers is that you can orient them toward a camera. After Effects is not a true 3D program and layers in 3D space are flat. When you are animating a camera, move through layers in 3D space, select your layers , and choose Layer > Transform > Auto Orient or press Cmd+Option+O (Ctrl+Alt+O). Choose Orient Towards Camera to have your layers turn as the camera moves through them. This gives the illusion that the layers have some dimension.