As an object moves along its path, you have two choices. Either the object maintains its orientation, or the object rotates to follow the path . Think of this animation as similar to a roller coaster car on its track.
Select a layer for which you have already created position keyframes.
Choose Layer>Transform>Auto Orient or press Cmd+Option+O (Ctrl+Alt+O).
Choose Orient Along Path, and to deactivate it choose Off.
You will find this incredibly useful when you are animating insects , unidentified flying objects, bobsleds, or any other object that needs to follow a curved path.